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Role of Bone Marrow-Derived CCR1+ Cells in Colon Cancer Invasion and Metastasis,l of invasive CRC whose TGF-β family signaling is blocked, we have found that immature myeloid cells (iMCs) are recruited from the bone marrow to the tumor invasion front. These iMCs express the matrix metalloproteinases MMP9 and MMP2 and the CC-chemokine receptor 1 (CCR1), and migrate towards the C