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Submitted on: 17 March 2008.
Revised on: 11 May 2008.
Accepted on: 25 June 2008.
___________________ANTARCTIC SCIENCE---CAMBRIDGE UNIV PRESS
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Submitted on: 21 June 2011.
Revised on: 06 October 2011.
Accepted on: 29 November 2011.
___________________ANTARCTIC SCIENCE---CAMBRIDGE UNIV PRESS
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